Research and Educational Publications from Shoals Marine Laboratory

This list includes publications based on work conducted at the Isles of Shoals using facilities and vessels of Shoal Marine Laboratory. Some publications include SML contribution numbers in their acknowledgments. Contribution numbers will be assigned after a paper is accepted for publication. To obtain an SML contribution number, email

To view SML's Publications List as PDF, click here.

Asterisks (*) denote an author who was an REU, RIFS, or other udergraduate student research intern at SML.

Scroll to bottom for Undergraduate, Masters, and Ph.D. theses conducted at Shoals Marine Laboratory.

Leigh, K.L.*, Sparks, J.P., and Bemis, W.E. 2016. Food Preferences of Atlantic Hagfish, Myxine glutinosa, Assessed by Experimental Baiting of Traps. Copeia 104(3):623-627.

Jacobson, S., J. Seavey and R. Mueller. 2016. Integrated science and art education for creative climate change communication. Ecol. and Soc. 21(3):30.

Frederick, P., N. Vitale, B. Pine, J. Seavey, and L. Sturmer. 2016. Reversing A Rapid Decline In Oyster Reefs: Effects Of Durable Substrate On Oyster Populations, Elevations, And Aquatic Bird Community Composition. J Shellfish Res 35: 359-367.

Byers, J.E., Z.C. Holmes and A.M.H. Blakeslee. 2016.  Consistency of trematode infection prevalence in host populations across large spatial and temporal scales.  Ecology. Ecology  97.7 (2016): 1643-1649. (SML Contribution 185).

Farina, S.C. and W.E. Bemis. 2016. Functional morphology of gill ventilation of the goosefish, Lophius americanus (Lophiiformes: Lophiidae)Zoology. Available online 24 February 2016, ISSN 0944-2006,

Bressman N.R.*, S.C. Farina, and A.C. Gibb. 2016. Look before you leap: Visual navigation and terrestrial locomotion of the intertidal killifish Fundulus heteroclitus. J. Exp. Zool. 325A:57–64.

Gough, W.T., S.C. Farina, F.E. Fish. 2015. Aquatic burst locomotion by hydroplaning and paddling in common eiders (Somateria mollissima). J. Exp. Biol. 218:1632-1638; doi: 10.1242/jeb.114140.

Bonter, D.N., M.C. Moglia* and L.E. DeFisher*. 2015. Sons don’t take advantage of a head start: Parity in offspring sex ratios despite greater initial investment in males. J. Avian Biol. 46:1-8. (SML Contribution 177)

Keogh, C.L., M.E. Sanderson*, and J.E. Byers. 2016. Local adaptation to parasite selective pressure: comparing three congeneric co-occurring hosts. Oecologia. 180:137-147. (SML Contribution 171)

Moisander, P.H., A.D. Sexton, and M.C. Daley. 2015. Stable associations masked by temporal variability in the marine copepod microbiome. PLoS ONE 10: e0138967. doi:10.1371/journal.pone.0138967. (SML Contribution 184)

Covino, K.M, S.R. Morris, and F.R. Moore. 2015. Patterns of testosterone in three Nearctic–Neotropical migratory songbirds during spring passage. Gen. Compar. Endro. 224:186-193. (SML Contribution 182)

Covino, K.M., 2015. Determination of sex using morphometrics in the northern waterthrush (Parkesia noveboracensis) and Swainson’s thrush (Catharus ustulatus). Wilson J. Ornithol. 127:706-711. (SML Contribution 180)

Shah, S.S*., E.I. Greig, S.A. MacLean*, and D.N. Bonter. 2015. Risk-based alarm calling in a nonpasserine bird. Anim. Behav. 106:129-136. (SML Contribution 166)

Bonter, D.N., S.A. MacLean*, S.S. Shah*, and M.C. Moglia*. 2014. Storm induced shifts in optimal nesting sites: A potential effect of climate change. J. Ornithol. 155: 631-638.  (SML Contribution 167)

Ellis, J.C., S.M. Bogdanowicz, M.C. Stoddard, and L.W. Clark. 2014. Hybridization of a lesser black-backed gull and herring gulls in eastern North America. Wilson J. Ornithol. 126: 338-345. (SML Contribution 168)

Garrett, J.N. and F.E. Fish. 2014. Kinematics of terrestrial locomotion in harbor seals and gray seals: Importance of spinal flexion by amphibious phocids. Mar. Mammal Sci. 32:  459-478.  DOI: 10.111/mms.2170

Mychajliw*, A.M. and R.G. Harrison. 2014. Genetics reveal the origin and timing of a cryptic insular introduction of muskrats in North America.  PLoS ONE. doi: 10.1371/journal.pone.0111856

Dijkstra, J.A., W.J. Lambert, L.G. Harris. 2013. Introduced species provide a novel temporal resource that facilitates native predator population growth. Bio. Invasions. 15:911-919.

Dorais, M.J. 2013. The Appledore Island Diorite of the Rye Complex, Cpastal New Hampshire and Maine: Determining the Identity of an Enigmatic Terrane. Abstr. with Programs 45: 107.

MacLean*, S.A. and D.N. Bonter. 2013.  The sound of danger: threat sensitivity to predator vocalizations, alarm calls, and novelty in gulls. PLoS ONE. 8(12): e82384. doi:10.1371/journal.pone.0082384. (SML Contribution 166)

DeFisher*, L.E. and D.N. Bonter. 2013. Effects of invasive European fire ants (Myrmica rubra) on Herring Gull (Larus argentatus) reproduction. PLoS ONE 8(5):e64185. (SML Contribution 163)

Simonis, J.L. 2013. Predator ontogeny determines trophic cascade strength in freshwater rock pools. Ecosphere 4: art62. (SML Contribution 165)

Brown, JM, NR Felice, NB Scalfone, and I Hewson. 2012. Influence of habitat confluence on aquatic microbial assemblages in experimental mesocosms. Aquat. Microb. Ecol. 66: 33-40.  (SML Contribution 160)

Simonis, J.L., D. Neuharth-Keusch*, and I. Hewson. 2012. Aquatic bacterial assemblage variability in the supra littoral zone of Appledore Island, Gulf of Maine. FEMS Microb. Ecol. 80: 501-508.

Ellis, J.C., K.E. Allen, M.S. Rome and M.J. Shulman. 2012. Choosing among mobile prey species:  Why do gulls prefer a rare subtidal crab over a highly abundant intertidal one? J. Exp. Mar. Biol. Ecol. 416-417: 84-91. (SML Contribution 158)

Seeley, R.H. and W.H. Schlesinger. 2012. Sustainable seaweed cutting? The rockweed (Ascophyllum nodosum) industry of Maine and the Maritime provinces. Ann. NY Acad. Sci. 1249: 84-103.

Seeley, T.D., P. K. Visscher, T. Schlegel, P. M. Hogan, N. R. Franks and J. A. R. Marshall. 2012. Stop signals provide cross inhibition in collective decision-making by honeybee swarms. Science. 335: 108-111.

Suomala, R.W., S.R. Morris, and K.J. Babbitt. 2012. Comparison of migrant songbird stopover ecology on two islands in the Gulf of Maine. Wilson J. Ornithol. 124:217-229. (SML Contribution 154)

Dijkstra, J.A., R. Nolan. 2011. Potential of the invasive colonial ascidian, Didemnum vexillum, to limit the escape response of the sea scallop, Placopecten magellanicus. Aquatic Invasions. 6(4): 451-456.

Hewson, I., J.M. Brown, S.A. Gitlin and D.F. Doud.  2011. Nucleopolyhedrovirus detection and distribution in terrestrial, freshwater and marine habitats of Appledore Island, Gulf of Maine. Microb. Ecol. 62: 48-57, doi: 10.1007/s00248-9856-1. (SML Contribution 156)

Elias, S.P., R. P. Smith, S.R. Morris, P.W. Rand, C. Lubelczyk and E.H. Lacombe. 2011. Density of Ixodes scapularis ticks on Monhegan Island after complete deer removal: A question of avian importation?  J. Vector Ecol.  36: 11-23, doi: 10.1111/j.1948-7134.2011.00136.x. (SML Contribution 155)

Martiny, A.C., J.B.H. Martiny, C. Weihe, A. Field and J.C. Ellis. 2011. Functional metagenomics reveals previously unrecognized diversity of antibiotic resistance genes in gulls. Front. Microbiol. 2: doi: 10.3389/fmicb.2011.00238.

Savoca*, M.S., D.N. Bonter, B. Zuckerberg, J.L. Dickinson, and J.C. Ellis. 2011. Nesting density is an important factor affecting chick growth and survival in the herring gull. The Condor 113: 565-571. (SML Contribution 157)

Suomala, R.W., S.R. Morris, K.J. Babbitt, and T.D. Lee. 2010. Migrant songbird species distribution and habitat use during stopover on two islands in the Gulf of Maine. Wilson J. Ornithol. 122:725-737.  (SML Contribution 153)

Seeley, T.D.  2010.  Honeybee Democracy.  Princeton, N.J.  Princeton Univ. Press.

Rangel, J., S.R. Griffin, and T.D. Seeley. 2010.  An oligarchy of nest-site scouts triggers a honeybee swarm’s departure from the hive.  Behav. Ecol. Sociobiol.  66: 979-987.

Rangel, J., S.R. Griffin, and T.D. Seeley. 2010.  Nest-site defense by competing honey bee swarms during house-hunting.  Ethology 116: 608-618.

Mulder, C.P.H. and J.C. Ellis. 2010. Seabird Island Ecology. In Encyclopedia of Life Sciences (ELS). John Wiley & Sons, Ltd: Chichester. Doi: 10.1002/9780470015902.a0022557.

Dantzker, M. and D.O. Brown. 2009. Signals for Survival (High-definition video). Sinauer, Sunderland, Massachusetts. 38 minutes.

List, C., C. Elsholtz, and T.D. Seeley.  2009.  Independence and interdependence in collective decision making:  an agent-based model of nest-site choice by honeybee swarms.  Philos. T. R. Soc. Lon. B 364: 755-762.

Freeman, A., J. Meszaros*, and J.E. Byers. 2009. Poor phenotypic integration of blue mussel inducible defenses in multiple predator environments. Oikos 118: 758-766.

Perez, K.O., R.L. Carlson, M.J. Shulman, J.C. Ellis. 2009. Why are intertidal snails rare in the subtidal? Predation, growth and the vertical distribution of Littorina littorea (L.) in the Gulf of Maine. J. Exp. Mar. Biol. Ecol. 369: 79-86.

Blakeslee*, A. and J.E. Byers. 2008. Using parasites to inform ecological history: comparisons among three congeneric marine snails. Ecology 89: 1068-1078.

Blakeslee*, A., J.E. Byers, M.P. Lesser. 2008. Solving cryptogenic histories using host and parasite molecular genetics: the resolution of Littorina littorea's North American origin. Mol. Ecol. 17: 3684-3696.

Bogomolni, A., R. Gast, J.C. Ellis, M. Dennett, K. Pugliares, B. Lentell, M. Moore. 2008. Victims or vectors: a survey of vertebrate zoonoses from coastal waters of the Northwest Atlantic. Dis. Aquat. Organ. 81: 13-38.

Byers, J.E., A. Blakeslee*, E. Linder, A. Cooper, and T. Maguire*. 2008. Controls of spatial variation in the prevalence of trematode parasites infecting a marine snail. Ecology 89: 439-451.

Ellis, J.C., M.C. Stoddard*, and L.W. Clark.  2008. Breeding by a Lesser Black-backed Gull (Larus fuscus) on the Atlantic coast of North America. N. Am. Birds. 61: 546-548. (SML Contribution  144)

Griffen, B.D. and T. Williamson.  2008. Influence of predator density on nonindependent effects of multiple predator species. Oecologia. 155: 151-159.

Nichols, W.F. and V.C. Nichols. 2008. The land use history, flora, and natural communities of the Isles of Shoals, Rye, New Hampshire and Kittery, Maine. Rhodora 110: 245-295.

Passino, K.M., T.D. Seeley, and P.K. Visscher. 2008. Swarm cognition in honey bees. Behav. Ecol. Sociobiol. 62: 401-414.

Rangel, J. and T.D. Seeley. 2008. The signals initiating the mass exodus of a honey bee swarm from its nest. Anim. Behav. 76: 1943-1952.

Rittschof*, C.C. and T.D. Seeley. 2008. The buzz-run: how honey bees signal ‘Time to go!” Anim. Behav. 75: 179-187.

Schultz, K., K.M. Passino, and T.D. Seeley. 2008. The mechanism of flight guidance in honeybee swarms: subtle guides or streaker bees? J. Exp. Biol. 211: 3287-3295.

Seeley, T.D. and P.K. Visscher. 2008. Sensory coding of nest-site value in honeybee swarms. J. Exp. Biol. 211: 3691-3697.

Dijkstra, J., L. Harris and E. Westerman. 2007. The long-term distribution and ecology of four invasive colonial ascidians in the Gulf of Maine. J. Exp. Mar. Biol. and Ecol. 342: 61-68.

Ellis, J.C., M.J. Shulman, H. Jessop, R. Suomala*, S. Morris, V. Seng, M. Wagner, and K. Mach*.  2007. Impact of a few mammalian predators on breeding success in large colonies of Herring Gulls and Great Black-backed Gulls. Waterbirds. 30: 375-383.

Ellis, J.C., M.J. Shulman, M. Wood, J.D. Witman, S. Lozyniak*. 2007. Regulation of cross-ecosystem food webs by avian predators on New England rocky shores. Ecology. 88: 853-863.

Visscher, P.K. and T.D. Seeley. 2007. Coordinating a group departure: who produces the piping signals on honeybee swarms? Behav. Ecol. Sociobiol. 61: 1615- 1621.

Wood, C.L.*, J.E. Byers, K. Cottingham, I. Altman, M. Donahue, and A. Blakeslee*. 2007. Parasites alter community structure. Proc. Nat. Acad. Sci. USA. 104: 9335-9339.

Beekman, M., R.L. *Fathke, and T.D. Seeley. 2006. How does an informed minority of scouts guide a honey bee swarm as it flies to its new home? Anim. Behav. 71: 161-171.

Carlson*, R.L, M.J. Shulman, J.C. Ellis.  2006.  Factors contributing to spatial heterogeneity in the abundance of the common periwinkle Littorina littorea (L.)  J. Mollus. Stud. 72: 149-156. (SML Contribution 124)

Ellis, J.C., J.M. Fariña, and J.D. Witman. 2006.  Nutrient transfer from sea to land: the case of gulls and cormorants in the Gulf of Maine.  J. Anim. Ecol. 75: 65-574.

Ellis, J.C. and T.P. Good. 2006.  Nest attributes, aggression, and breeding success of gulls in single and mixed-species subcolonies.  The Condor. 108: 211-219.

Fischer, E., A. Pszenny, W. Keene, J. Maben, A. Smith, A. Stohl, and R. Talbot. 2006. Nitric acid phase partitioning and cycling in the New England coastal atmosphere. J. Geophys. Res. 111, D23S09, doi:10.1029/2006JD007328. (SML Contribution 126)

Jones, P.L. and M.J. Shulman. 2006. Subtidal-intertidal trophic links:  American lobsters [Homarus americanus (Milne-Edwards)] forage in the intertidal zone on nocturnal high tides. J. Exp. Mar. Biol. Ecol. 361: 98-103.

Morris, S. R., A. M. Larracuente, K. M. Covino*, M. S. Mustillo, K. E. Mattern, D. A. Liebner, and H. D. Sheets.  2006.  Utility of open population models: Limitations posed by parameter estimability in the study of migratory stopover.  Wilson J. Ornithol. 118: 513-526. (SML Contribution 127)

Passino, K. and T.D. Seeley. 2006. Modeling and analysis of nest-site selection by honey bee swarms: the speed and accuracy trade-off. Behav. Ecol. Sociobiol. 59: 427-442.

Seeley, T.D., P.K. Visscher, and K.M. Passino. 2006. Group decision making in honey bee swarms. Amer. Sci. 94: 220-229.

Ellis, J.C., W. Chen*, B. O'Keefe*, M.J. Shulman, J.D. Witman. 2005. Gull predation on crabs in the rocky intertidal and shallow subtidal zones of coastal New EnglandJ. Exp. Mar. Biol. Ecol. 324: 31-43. (SML Contribution 120)

Morris, S. R., D. A. Liebner, A. M. Larracuente, E. M. Escamilla, and H. D. Sheets.  2005.  Multiple-day constancy as an alternative to pooling for estimating mark-recapture stopover length in Nearctic-Neotropical migrant landbirds.  Auk 122: 319-328.  (SML Contribution 117)

Morris, S. R., E. M. Turner, D. A. Liebner, A. M. Larracuente, and H. D. Sheets.  2005.  Problems associated with pooling mark-recapture data prior to estimating stopover length for migratory passerines. Pages 673-679 in Bird Conservation Implementation and Integration in the Americas: Proc. Third Inter Partners in Flight Conference 2002 (C.J. Ralph and T.D. Rich, eds.). U.S.D.A. Forest Service, PSW-GTR-191, Albany, CA. (SML Contribution 109)

Sive, B. C., Y. Zhou, D. Troop, Y. Wang, W. C. Little, O. W. Wingenter, R. S. Russo, R. K. Varner, and R. Talbot. 2005. Development of a cryogen-free concentration system for measurements of volatile organic compounds, Anal. Chem. 77: 6989-6998,10.1021/ac0506231. (SML Contribution 122)

Stefaniak*, L.M., J. McAtee and M.J. Shulman. 2005. The costs of being bored: effects of a clionid sponge on the gastropod Littorina littorea.  J. Exp. Mar. Biol. Ecol. 327: 103-114.  (SML Contribution 121)

Cully, S.M.* and T.D. Seeley. 2004. Self-assemblage formation in a social insect: the protective curtain of a honey bee swarm. Insectes Sociaux 51: 317-324.

Novak, M.* 2004. Diurnal activity in a group of Gulf of Maine decapods. Crustaceana 77(Part 5): 603-620. (SML Contribution 118)

Rome, M.S.* and J.C. Ellis. 2004. Foraging ecology and interactions between Herring Gulls and Great Black-backed Gulls in New England. Waterbirds. 27: 200-210. (SML Contribution 119)

Seeley, T.D. and P.K. Visscher. 2004. Group decision making in nest-site selection by honey bees. Apidologie 35: 1-16.

Seeley, T.D. and P.K. Visscher. 2004. Quorum sensing during nest-site selection by honeybee swarms. Behav. Ecol. Sociobiol. 56: 594-601.

Apprill*, A.M.  and M.P. Lesser. 2003. Effects of ultraviolet radiation on Laminaria saccharina in relation to depth and tidal height in the Gulf of Maine. Mar. Ecol. Prog. Ser. 256: 75-85. (SML Contribution 108)

Byrnes*, J. and J. Witman. 2003. Impact assessment of an invasive flatworm, Convoluta convoluta, in the southern Gulf of Maine. J. Exp. Mar. Biol. Ecol. 293: 173-191. (SML Contribution 132)

Morris, S. R., C. R. Pusateri and K. A. Battaglia.  2003.  Spring migration and stopover ecology of Common Yellowthroats (Geothlypis trichas) on Appledore Island, MaineWilson Bull. 115: 64-72. (SML Contribution 110)

Seeley, T.D. and P.K. Visscher. 2003. Choosing a home: how the scouts in a honey bee swarm perceive the completion of their group decision making. Behav. Ecol. Sociobiol. 54: 511-520.

Britton, N.F., S.C. Pratt, N.R. Franks, and T.D. Seeley. 2002. Deciding on a new home: how do honey-bees agree? Proc. R. Soc. Lon. B 269: 1383-1388.

Flesher, D. and F.H. Martini. 2002. Lampreys, Pages 16-19 in Bigelow and Schroeder’s Fishes of the Gulf of Maine, 3rd Edition. (B.B. Collette and G. Klein-MacPhee, eds.) Smithsonian Institute Press, Washington, D. C.

Martini, F.H. and D. Flescher. 2002. Hagfishes, Pages 9-16 in Bigelow and Schroeder, Fishes of the Gulf of Maine, 3rd Edition. (B.B. Collette and G. Klein-MacPhee, eds.) Smithsonian Institute Press, Washington, D. C.

Wheeler, A.G. Jr. 2002. Chilacis typhae (Perrin) and Holcocranum saturejae (Kolenati) (Hemiptera: Lygaeoidea: Artheneidae): Updated North American distributions of two palearctic cattail bugs. 2002. Proc. Entomol. Soc. Wash. 104: 24-32.

Gellin, C. E. and S. R. Morris. 2001. Patterns of movement during passerine migration on an island stopover site. Northeast. Nat. 8: 253-266. (SML Contribution 107)

Morris, S.R. and J. Glasgow. 2001. Spring and fall migration of American Redstarts on Appledore Island, Maine. Wilson Bull. 113: 202-210. (SML Contribution 106)

Seeley, T.D. and S.C. Buhrman. 2001. Nest-site selection in honey bees: How well do swarm implement the "best-of-N" decision rule? Behav. Ecol. Sociobiol. 49: 416-427. (SML Contribution 105)

Levin, P.S., J.A. Coyer, R. Petrik, and T.P. Good. 2002. Community-wide effects of non-indigenous species on temperate rocky reefs. Ecology. 83: 3182-3193. (SML Contribution 104)

Weidenmiller, A., and T.D. Seeley. 1999. Imprecision in the waggle dances of honey bees for nearby food sources: Error or adaptation? Behav. Ecol. and Sociobiol. 46: 190-199. (SML Contribution 103)

Morris, S.R., and M.T. Bradley. 2000. Is tail feather shape a reliable indicator of age in warblers and thrushes? N. Amer. Bird Bander  25: 125-131. (SML Contribution 102)

Morris, S. R., A. J. Donovan, and S. M. Agugliaro. 2003. Accuracy of sex determination of hatch-year Common Yellowthroats (Geothlypis trichas) during the fall. N. Amer. Bird Bander  28: 105-110. (SML Contribution 102)

Genovese, S.J and J.D. Witman 1999. Interactive effects of flow speed and particle concentration on growth rates of an active suspension feeder. Limnol. Oceanogr. 44: 1120-1131. (SML Contribution 101)

Rivest, B., J. Coyer and S. Tyler. 1999. The first known invasion of a free-living marine flatworm. Biol. Invasions 1: 393-394. (SML Contribution 100)

Martini, F. H. 1998. The Ecology of Hagfishes, in The Biology of Hagfishes, J. M. Joergenson ed., Chapman and Hall Pubs., London, pp: 57-77.

Martini, F.H. 1998. Secrets of the Slime Hagfish. Sci. Amer. (October 1998) 228: 44-49.

Martini, F.H., J.B. Heiser, and M.P. Lesser. 1998. A population profile for Atlantic Hagfish, Myxine glutinosa l., in the Gulf of Maine, part 2: Morphological variation in populations of Myxine glutinosa l. in the North Atlantic ocean. Fish. Bull. 96: 516-514.

Levin, P.S., W. Chiasson, J.M. Green. 1997. Geographic differences in recruitment and population structure of a temperate reef fish. Mar. Ecol. Pro. Ser. 161: 23-35.

Martini, F., J.B. Heiser, and M.P. Lesser. 1997. Ecology of the Hagfish, Myxine glutinosa l., in the Gulf of Maine II: Potential impact on benthic communities and commercial fisheries. J. Exp. Mar. Biol. Ecol. 214: 97-106

Lesser, M.P., E.H. Martini, and J.B. Heiser. 1996. Ecology of the Hagfish, Myxine glutinosa l., in the Gulf of Maine I: Metabolic rates and energetics. J. Exp. Mar. Biol. Ecol. 208: 215-225.

Levin, P.S. 1996. Recruitment in a temperate demersal fish: Does larval supply matter? Limnol. Oceanogr. 41: 672-679.

Martini, F.H., J.B. Heiser, and M.P. Lesser. 1996. A population profile for Atlantic Hagfish, Myxine glutinosa l., in the Gulf of Maine, part 1: Morphometrics and reproductive state. Fish. Bull. 95: 311-320.

Morris, S.R. 1996. Mass loss and probability of stopover by migrant warblers during spring and fall migration. J. Field Ornithol. 67: 456-462.

Morris, S.R., D.W. Holmes, and M.E. Richmond. 1996. A ten-year study of the stopover patterns of migratory passerines during fall migration on Appledore Island, Maine. The Condor. 98: 395-409.

Smith, R.P., Jr., P.W. Rand, E. H. Lacombe, S.R. Morris, D.W. Holmes, and D.A. Caporale. 1996. Role of bird migration in the long distance dispersal of Ixodes dammini, the vector of lyme disease. J. Infect. Dis. 174: 221-224. (SML Contribution 133).

Levin, P.S. 1994. Small-scale recruitment variation in a temperate fish: the roles of macrophytes and food supply. Environ. Biol. Fish. 40: 271-281.

Levin, P.S. 1994. Fine-scale variation in recruitment of a temperate demersal fish: the importance of settlement versus post-settlement loss. Oecologia. 97: 124-133.

Morris, S.R., M.E. Richmond, and D.W. Holmes. 1994. Patterns of stopover by warblers during spring and fall migration on Appledore Island, Maine. Wilson Bull. 106: 703-718.

Factor, J.R., and B.L. Dexter. 1993. Suspension feeding in larval crabs (Carcinus maenas). J.  Mar. Biol. Assoc. U.K. 73: 207-211.

Levin, P.S. 1993. Habitat structure, conspecific presence and spatial variation in the recruitment of a temperate reef fish. Oecologia 94: 176-185.

Witman, J.D., and K.P. Sebens. 1992. Regional variation in fish predation intensity: a historical perspective in the Gulf of Maine. Oecologia 90: 305-315.

Kingsbury, J.M.  1991. Here’s How We’ll Do It: An Informal History of the Construction of the Shoals Marine Laboratory, Appledore Island, Maine. Bullbrier Press, Ithaca, NY. 364 pp.

Levin, P.S. and A.C. Mathieson 1991. Variation in a host-epiphyte relationship along a wave exposure gradient. Mar. Ecol. Prog. Ser. 77: 271-278.

Witman, J.D. 1987. Subtidal coexistance: storms, grazing, mutualism, and the zonation of kelps and mussels. Ecol. Monogr. 57: 167-187.

Seeley, R. H. 1986. Intense natural selection caused a rapid morphological transition in a living marine snail. Proc. Natl. Acad. Sci. USA. 83: 6897-6901.

Witman, J.D. 1985. Refuges, biological disturbance, and rocky subtidal community structure in New England. Ecol. Monogr. 55: 421-445.

Seeley, T.D. 1983. Division of labor between scouts and recruits in honeybee foraging. Behav. Ecol. Sociobiol. 12: 253-259.

Seeley, T.D. 1982. How honeybees find a home. Sci. Amer. 247: 158-168.

Boden, G.T. 1979. The effect of depth on summer growth of Laminaria saccharina (Phaeophyta, Laminariales). Phycologia 18: 405-408.

Seeley, T.D., R.A. Morse, and P.K. Visscher. 1979. The natural history of the flight of honey bee swarms. Psyche 86: 103-113.

Seeley, T.D. 1977. Measurement of nest cavity volume by the honey bee (Apis mellifera). Behav. Ecol. Sociobiol. 2: 201-227.

Boden, G.T. 1977.  The Vascular Flora of Appledore Island. Shoals Marine Laboratory, Ithaca, NY. 46 pp.

Gowen, A.G. and H.F. Mulligan. 1978.  A Photographic Guide to the Phytoplankton of the Coastal Waters, Gulf of Maine. Shoals Marine Laboratory, Ithaca, NY. 164 pp.

Fowler-Billings, Katharine. 1977. Geology of the Isles of Shoals. Published by the State of New Hampshire Department of Resources and Economic Development. Concord, NH.

Kingsbury, J.M. 1975. Oil and Water: The New Hampshire Story. Shoals Marine Laboratory, Ithaca, NY. 102 pp.

Undergraduate Honors Theses resulting from SML REU and RIFS research

Brown, E. 2001. The role of food availability in determining the distribution and abundance of the sea star (Asterias vulgaris) and the green sea urchin (Strongylocentrotus droebachiensis). Honor’s Thesis. Brown University.

Byrnes, J. 2001. The potential impact of an invasive acoel, Convoluta convoluta, in the Gulf of Maine. Honor’s Thesis. Brown University.

Chen, W. 2001. Avian predation in the intertidal: Impact of the Great Black-backed Gull, Larus marinus (Linnaeus), on crab abundance. Honor’s Thesis. Cornell University.

Kim, A. 2001. Encapsulation as protection from ultraviolet radiation for larval stages of the intertidal gastropod, Nucella lapillus. Honor’s Thesis. Smith College.

Kraft, N. 2001. The role of decapod predation in limiting the lower distribution of the intertidal snail Littorina littorea in the Gulf of Maine. Honor’s Thesis. Brown University.

Masters Theses & Ph.D. Dissertations

Simonis, J. 2012. (Cornell University) Ph.D. Dissertation

Burbidge, S. 2008. (Antioch New England) Thesis, Master of Science. Nest-site Selection and Response to Habitat Manipulation by Roseate Terns (Sterna dougallii) on Seavey Island, New Hampshire.

Morris, S.R. 1996. (Cornell University) Ph.D. Dissertation. The migration and stopover ecology of neotropical migrants on Appledore Island, Maine.

Morris, S.R. 1993. (Cornell University) Thesis, Master of Science. Patterns of stopover migratory passerines on Appledore Island, Maine: An analyses of banding records from 1983-1991.

Lyman, E. G. 1988. (UNH) Thesis, Master of Science. A Comparison of Island and Mainland Populations of the Muskrat, Ondatra zibethicus.

Doherty, K  1986. (UNH) Thesis, Master of Science. An evaluation of secondary wastewater treatment alternatives for saline domestic wastewater.

Montagnes, D. 1986. (UNH) The annual cycle of planktonic ciliates in the waters surrounding the Isles of Shoals, Gulf of Maine: Estimates of biomass and production.

McGill-Harelstad, P. A. 1985. (Cornell University)  Ph.D. Dissertation. Mechanisms and Consequences of Interspecific Interactions among Gulls. 113 pp.

Weeks, H.J. 1985. (Cornell University)  Ph.D. Dissertation. Ecology of the Three Spined Stickleback (Gasterosteus aculeatus): Reproduction in rocky tidepools.

Lewis, S.J. 1982. (Cornell University) Ph.D. Dissertation. Effects of oil on avian productivity and population dynamics. 172 pp.

Boden, G.T. (Cornell University) 1977. Thesis, Master of Science. Some aspects of the ecology and growth of the brown seaweeds (Laminaria spp., Agarum spp., and Alaria spp.) in the Southern Gulf of Maine.

McGill, P.A. 1977. (Cornell University) Thesis, Master of Science. Breeding Ecology and Competition between Great Black-backed and Herring Gulls. 88 pages.

Kinner, Nancy 1976. (UNH) Thesis, Master of Science. An evaluation of the treatment of saline domestic wastewater using rotating biological contactor.

Martini, R.F. 1975. (Cornell University) PhD dissertation. Product development and nutritional evaluation of underutilized species of sharks.